By Ralf Heinrich
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40C). However, it is 45 clear from molecular analysis that the South American forms are comprised of at least three cryptic taxa, each separate from the North American one, and that the European form is distinct at the species level from the North American one. With two exceptions, the remaining species are restricted to the Americas. D. párvula and D. ambigua are widespread in temperate and montane regions of South America and north temperate USA. D. retrocurva, D. neo-obtusa, D. minnehaha are restricted to north temperate USA.
D. atkinsoni, D. bolivari and D. triquetra are considered close because they share an anteriorly expanded dorsal ridge with varying degrees of spination. They have been linked to another closely related set of species D. mediterránea, D. hispánica, D. dolichocephala and D. deserti, through the charac ters of their trunk limbs, and these may be related to the two Asian species D. fusca and D. tibetana. Another major group is the D. carinata complex with associated species D. cephalata and D. nivalis, which appears to be more distantly related to the D.
5% (Schwenk & Spaak 1997; Giessler et al. 1999). 2 The Daphnia pulex species complex The other major species group that is known to have high degrees of interspecies hybrid ization and which has been the subject of intense molecular investigation is the D. pulex complex (Hebert & Loaring 1986; Hebert 1987; Cerny & Hebert 1993; Hebertet al. 1989, 1993; Dufresne & Hebert 1994, 1995, 1997; Ward et al. 1994; Weider & Hobaek 1994, 1997, 2003; Van Raay & Crease 1995; Weider et al. 1996; Crease et al.